We are pleased to announce the recipient of the second annual SSE Lifetime Achievement Award [1], Dr. Richard E. Lenski. The Lifetime Achievement Award was created to recognize individuals who have made substantial contributions to the study of evolution, who have demonstrated outstanding mentorship of trainees, and/or who have provided noteworthy service to the evolution community. Dr. Lenski is the John Hannah Distinguished Professor of Microbial Ecology at Michigan State University (MSU). For more than 30 years, he has directed one of the most significant experiments in evolutionary biology, the E. coli Long Term Evolution Experiment [2] (LTEE). He is one of the founding members of the BEACON Center for the Study of Evolution in Action at MSU [3], and today serves on its executive committee. Dr. Lenski has previously received a Guggenheim Fellowship and a MacArthur Fellowship, and has been elected to the American Academy of Microbiology, the European Molecular Biology Organization, the American Academy of Arts and Sciences, and the National Academy of Sciences. In addition to his outstanding research contributions, he is known for being an exceptional mentor and role model. He also served the community as President of SSE in 2013. Congratulations, Dr. Lenski!